Le genre Dryopteris 4/4

De P à Z

P/
Dryopteris paleolata (Pic.Serm.) Li Bing Zhang, 2012
Dryopteris pallida (Bory) Maire & Petitm., 1908
⤷ pallida subsp. balearica (Litard.) Fraser-Jenk., 1977
⤷ pallida subsp. libanotica (Rosenst.) E.Nardi, 1977
⤷ pallida subsp. raddeana (Fomin) E.Nardi, 1977

Dryopteris panda (C.B.Clarke) C.Chr., 1934
Dryopteris paomowanensis Ching & Z.Y.Liu, 1984
Dryopteris papuae-novae-guineae Li Bing Zhang, 2012
Dryopteris papuana C.Chr., 1905
Dryopteris parabissetiana H.J.Wei & Z.Y.Zuo, 2024
Dryopteris patula (Sw.) Underw., 1893
Dryopteris paucisora Copel., 1911
Dryopteris pauliae Fraser-Jenk., Widén & Gibby, 1997
Dryopteris peninsulae Kitag., 1935
Dryopteris pentheri (Krasser) C.Chr., 1905
Dryopteris peranema Li Bing Zhang, 2012
Dryopteris permagna M.G.Price, 1977
Dryopteris × picoensis Fraser-Jenk. & Gibby, 1981

    (D. affinis × D. intermedia subsp. azorica)

Dryopteris × pittsfordensis Sloss., 1904

    (D. carthusiana × D. marginalis)

Dryopteris podophylla (Hook.) Kuntze, 1891
Dryopteris poilanei Tardieu, 1941
Dryopteris polita Rosenst., 1914
Dryopteris polylepis (Franch. & Sav.) C.Chr., 1905
Dryopteris porosa Ching, 1938
Dryopteris protobissetiana K.Hori & N.Murak., 2015
Dryopteris × pseudoabbreviata Jermy, 1968

    (D. aemula × D. oreades)

Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, 2012
Dryopteris × pseudocommixta Sa.Kurata, 1964

    (D. commixta × D. kuratae)

Dryopteris pseudofilix-mas (Fée) Rothm., 1946
Dryopteris × pseudohangchowensis Miyam., 1984

    (D. hangchowensis × D. kuratae)

Dryopteris pseudolunanensis Tagawa, 1940
Dryopteris pseudoparasitica Alderw., 1924
Dryopteris pseudosieboldii Hayata, 1914
Dryopteris pseudosparsa Ching, 1938
Dryopteris pteridiiformis Christ, 1907
Dryopteris pulvinulifera (Bedd.) Kuntze, 1891
Dryopteris purpurascens Christ, 1907

Dryopteris pycnolepis Z.Y.Zuo & S.Y.Dong, 2025

Dryopteris pycnopteroides (Christ) C.Chr., 1913

R/

Dryopteris raiateensis (J.W.Moore) Li Bing Zhang, 2012
Dryopteris ramosa (C.Hope) C.Chr., 1905
Dryopteris × rarissima Sa.Kurata, 1964

    (D. commixta × D. sparsa)

Dryopteris raynalii Tardieu, 1965
Dryopteris redactopinnata Soumen K.Basu & Panigrahi, 1980
Dryopteris reflexosquamata Hayata, 1914
Dryopteris remota (A.Braun ex Doll) Druce, 1908
Dryopteris remotipinnula Bonap., 1925
Dryopteris renchangiana Z.Y.Zuo & D.Z.Li, 2021 publ. 2022
Dryopteris rhomboideo-ovata H.Itô, 1933
Dryopteris rodolfii J.P.Roux, 2004
Dryopteris × ronald-vianensis Pérez Carro & Fern.Areces, 2016

    (D. dilatata × D. guanchica)

Dryopteris rossii C.Chr., 1912
Dryopteris rosthornii (Diels) C.Chr., 1905
Dryopteris rubiginosa (Brack.) Kuntze, 1891
Dryopteris rubrobrunnea W.M.Chu, 1992
Dryopteris ruwenzoriensis C.Chr. ex Fraser-Jenk., 1986
Dryopteris ryo-itoana Sa.Kurata, 1967

S/

Dryopteris sabaei (Franch. & Sav.) C.Chr., 1905
Dryopteris sacrosancta Koidz., 1924
Dryopteris saffordii C.Chr., 1911
Dryopteris sandwicensis (Hook. & Arn.) C.Chr., 1905
Dryopteris × sardoa Fraser-Jenk., Reichst, 1975

    (D. oreades × D. tyrrhena)

Dryopteris × sarvelae Fraser-Jenk. & Jermy, 1977

    (D. carthusiana × D. expansa)

Dryopteris × satsumana Sa.Kurata, 1969

    (D. commixta × D. dickinsii)

Dryopteris saxifraga H.Itô, 1936

Dryopteris saxifragivaria Nakai, 1942
Dryopteris scabrosa (Kunze) Kuntze, 1891
Dryopteris schimperiana (Hochst.) C.Chr., 1905
Dryopteris schizopaleata Eb.Fisch. & Lobin, 2023
Dryopteris schnellii Tardieu, 1948
Dryopteris scottii (Bedd.) Ching, 1933
Dryopteris × separabilis Small, 1938

    (D. celsa × D. intermedia)

Dryopteris sericea C.Chr., 1913
Dryopteris serratodentata (Bedd.) Hayata, 1914
Dryopteris shiakeana H.Shang & Y.H.Yan, 2015
Dryopteris × shibipedis Sa.Kurata, 1966

    (D. kinkiensis × D. pacifica)

Dryopteris × shibisanensis Sa.Kurata, 1965

    (D. atrata × D. commixta)

Dryopteris shikokiana (Makino) C.Chr., 1905
Dryopteris shiroumensis Sa.Kurata & Nakam., 1960
Dryopteris sichotensis Kom., 1916
Dryopteris sieboldii (T.Moore) Kuntze, 1891
Dryopteris sikkimensis (Bedd.) Kuntze, 1891
Dryopteris simasakii (H.Itô) Sa.Kurata, 1970
Dryopteris sinonepalensis Z.Y.Zuo & Fraser-Jenk., 2021 publ. 2022

Dryopteris × sjoegrenii Fraser-Jenk., 1981

    (D. dilatata × D. intermedia subsp. azorica)

Dryopteris sledgei Fraser-Jenk., 1989
Dryopteris × slossoniae Wherry ex Lellinger, 1984

    (D. cristata × D. marginalis)

Dryopteris songyinghuensis J.X.Li & Xiao J.Li, 2025

Dryopteris sordidipes Tagawa, 1934
Dryopteris sororia (Maxon) M.A.McHenry, Sundue & Barrington, 2013

Dryopteris sparsa (D.Don) Kuntze, 1891

⤷ sparsa subsp. rectipinnula Fraser-Jenk., 2008

⤷ sparsa nothosubsp. lehalii Fraser-Jenk., 2008

   (D. sparsa subsp. rectipinnula × D. sparsa subsp. sparsa)

Dryopteris sphaeropteroides (Baker) C.Chr., 1905
Dryopteris splendens (Desv.) Kuntze, 1891
Dryopteris squamiseta (Hook.) Kuntze, 1891
Dryopteris stanley-walkeri Fraser-Jenk., 2001
Dryopteris stenolepis (Baker) C.Chr., 1905
Dryopteris stewartii Fraser-Jenk., 1979
Dryopteris subarborea (Baker) C.Chr., 1905
Dryopteris subatrata Tagawa, 1940
Dryopteris × subaustriaca Rothm., 1945

    (D. campyloptera × D. filix-mas)

Dryopteris subbipinnata W.H.Wagner & R.W.Hobdy, 1993
Dryopteris subcochleata Fraser-Jenk. & Khatri Chhetri, 2015
Dryopteris × subdiffracta H.J.Wei & Z.Y.Zuo, 2024

    (D. diffracta × D. polita)

Dryopteris subexaltata (Christ) C.Chr., 1905
Dryopteris subhikonensis K.Hori & N.Murak., 2018
Dryopteris subimpressa Loyal, 1969
Dryopteris sublacera Christ, 1909
Dryopteris submarginata Rosenst., 1914
Dryopteris subpycnopteroides Ching ex Fraser-Jenk., 1986
Dryopteris × subreflexipinna M.Ogata, 1935

    (D. diffracta × D. hasseltii)

Dryopteris subtriangularis (C.Hope) C.Chr., 1905
Dryopteris subtsushimensis K.Hori & N.Murak., 2018
Dryopteris × subuliginosa Charit., 2020

    (D. cristata × D. filix-mas)

Dryopteris × sugino-takaoi Sa.Kurata, 1965

    (D. lacera × D. polylepis)

Dryopteris sukungiana Z.Y.Zuo, 2022
Dryopteris sweetiorum Lorence & W.L.Wagner, 2011

T/

Dryopteris tahmingensis Ching, 1938
Dryopteris × takachihoensis Miyam., 1984

    (D. hangchowensis × D. uniformis)

Dryopteris takeuchiana Koidz., 1932
Dryopteris × telesii Fraser-Jenk., 1981

    (D. crispifolia × D. dilatata)

Dryopteris tenuicula C.G.Matthew & Christ, 1909
Dryopteris tenuipes (Rosenst.) Seriz., 1971
Dryopteris tetrapinnata W.H.Wagner & Hobdy, 1999
Dryopteris × tetsu-yamanakae Sa.Kurata, 1962

    (D. commixta × D. sieboldii)

Dryopteris × tiantangzhaiensis H.J.Wei & B.Chen, 2024

    (D. gymnophylla × D. tsoongii)

Dryopteris tingiensis Ching & S.K.Wu ex Fraser-Jenk., 1986
Dryopteris × tokudae Sugim., 1958

    (D. crassirhizoma × D. polylepis)

Dryopteris tokyoensis (Makino) C.Chr., 1905
Dryopteris × toyamae Tagawa, 1939

    (D. lacera × D. sieboldii)

Dryopteris transmorrisonensis (Hayata) Hayata, 1914
Dryopteris tricellularis J.P.Roux, 2002
Dryopteris × triploidea Wherry, 1960

    (D. carthusiana × D. intermedia)

Dryopteris tsoongii Ching, 1987
Dryopteris tsugiwoi Sa.Kurata, 1969
Dryopteris tsushimensis K.Hori & N.Murak., 2018
Dryopteris tsutsuiana Sa.Kurata, 1969
Dryopteris tyrrhena Fraser-Jenk. & Reichst, 1975

U/

Dryopteris × uliginosa (A.Braun ex Döll) Kuntze ex Druce, 1908

    (D. carthusiana × D. cristata)

Dryopteris undulata (Bedd.) Kuntze, 1891
Dryopteris unidentata (Hook. & Arn.) C.Chr., 1905

 ⤷ unidentata var. paleacea (Hillebr.) Herat ex Fraser-Jenk., 1994

Dryopteris uniformis (Makino) Makino, 1909
Dryopteris × uralensis Gureyeva & Moczalov, 2014

    (D. expansa × D. filix-mas)

V/

Dryopteris varia (L.) Kuntze, 1891
Dryopteris vescoi (Drake) C.Chr., 1905
Dryopteris × vidae Fraser-Jenk. & Gibby, 1980

    (D. pallida × D. vilarii)

Dryopteris vidyae Fraser-Jenk., 1989
Dryopteris villarii (Bellardi) Woyn., 1915
Dryopteris viscidula (Mett.) Kuntze, 1891

W/

Dryopteris wallichiana (Spreng.) Hyl., 1953

⤷ wallichiana subsp. convexa Fraser-Jenk., 2023

⤷ wallichiana subsp. coriacea (Fraser-Jenk.) Fraser-Jenk., 1994

⤷ wallichiana subsp. madagascariensis (C.Chr.) J.P.Roux, 2009
⤷ wallichiana subsp. madrasensis (Fraser-Jenk.) Fraser-Jenk., 1994
⤷ wallichiana subsp. nepalensis Fraser-Jenk., 1997

⤷ wallichiana subsp. pachyphylla Fraser-Jenk. & Ralf Knapp, 2018

⤷ wallichiana subsp. parrisiae (Fraser-Jenk.) Fraser-Jenk., 2022

⤷ wallichiana subsp. reichsteinii Fraser-Jenk., 1996
 ⤷ wallichiana f. rubiginosa (Fomin) Fraser-Jenk., 2024
 ⤷ wallichiana f. gigantea (Fraser-Jenk.) Fraser-Jenk., 2024

Dryopteris wardii (Baker) Kuntze, 1891
Dryopteris × watanabei Sa.Kurata, 1965

    (D. crassirhizoma × D. uniformis)

Dryopteris wattsii M.McKeown, Sundue & Barrington, 2012
Dryopteris × wechteriana Fraser-Jenk., 1989

    (D. chrysocoma × D. nigropaleacea)

Dryopteris whangshangensis Ching, 1938
Dryopteris wideniana Fraser-Jenk., 2015
Dryopteris woodsiisora Hayata, 1916
Dryopteris wulingshanensis J.P.Shu, Y.H.Yan & R.J.Wang, 2021
Dryopteris wuyishanica Ching & P.C.Chiu, 1987

X/

Dryopteris xanthomelas (Christ) C.Chr., 1913
Dryopteris xunwuensis Ching & K.H.Shing, 1990

Y/

Dryopteris yakusilvicola Sa.Kurata, 1957

Dryopteris × yamashitae Sa.Kurata, 1969

    (D. hayatae × D. sparsa)

Dryopteris yongdeensis W.M.Chu ex S.G.Lu, 1990
Dryopteris yoroii Seriz., 1971
Dryopteris × yuyamae Sa.Kurata, 1966

    (D. kuratae × D. uniformis)

Z/

Dryopteris zayuensis Ching & S.K.Wu, 1983
Dryopteris zhenfengensis P.S.Wang & X.Y.Wang, 2001
Dryopteris × zygoparentalis Fraser-Jenk., 1989

    (D. gamblei × D. scottii)




⏪

Le genre Dryopteris 1/4

Dryopteris Adans.

Première publication 1763, nom. cons.

Classifié pou certains dans la famille

des Polypodiaceae.

Étymologie :

Du grec, δρῦς drýs : chêne ;

mais aussi δόρυ, dóry : bois.

et de πτέρις ptéris : fougère.

Ce réfère au fait que les espèces du genre ont une forte prédilection

au zones boisées.

Synonyme :

Acrophorus C. Presl, 1836

Acrorumohra (H. Itô) H. Itô, 1939
Arthrobotrys Wall., 1829, not validly publ.
Desmopodium J.Sm., 1861

Diacalpe Blume, 1828

Dichasium (A. Braun) Fée, 1852

Diclisodon T. Moore, 1857

Dryopsis Holttum & P.J. Edwards, 1986

Filix Ség., 1754, nom. rej.
Filix-mas Hill ex Farw., 1931
Gymnothalamium Zenker ex Kunze, 1851, not validly publ.

Lophodium Newman, 1851

Nephrodium Marthe ex Michx., 1803

Nothoperanema (Tagawa) Ching, 1966

Peranema D. Don, 1825

Pycnopteris T. Moore, 1855

Revwattsia D.L. Jones, 1998
Sphaeropteris Wall., 1830, nom. illeg.

Nom de genre accepté :

                   /

Le genre est décrit comme ambigu par certains.

D'après certains organismes de classement,

le genre se diviserait en sous-genres et sections,

mais tous les sous-genres et sections ne sont pas légitimes

(voir la liste ci-dessous) :

Genre : Dryopteris Adans. 1763
              ↓   ⤷Sect. Acrophorus (C. Presl) Li Bing Zhang, 2012

              ↓   ⤷Sect. Acrorumohra (H. Itô) Li Bing Zhang & H. He Fl.,

              ↓  ↓           2013

              ↓   ⤷Sect. Aemulae Fraser-Jenk., 1986

              ↓   ⤷Sect. Amplae C. Chr., 1934

              ↓  ↓             ⤷Ser. Championianae Ching ex S.F. Lu, 1999

              ↓   ⤷Sect. Caespitosae S.G. Lu, 1990

              ↓   ⤷Sect. Chrysocomae S.G. Lu, 1990

               ⤷Sous-Genre : Ctenitis

               ⤷Sous-Genre : Cyclosorus

              ↓   ⤷Sect. Diacalpe (Blume) Li Bing Zhang, 2012

              ↓   ⤷Sect. Dryopsis (Holttum & P.J. Edwards) Li Bing Zhang,

              ↓  ↓          2012

              ↓   ⤷Sect. Dryopteris

               ⤷Sous-Genre : Dryopteris
              ↓    ⤷ sect. Erythrovariae H. Itô, 1936

               ⤷Sous-Genre : Erythrovariae (H. Itô) Fraser-Jenk., 1986

               ⤷Sous-Genre : Erythroviae Kraenzl., 1999

               ⤷Sous-Genre : Eudryopteris C. Chr., 1906

              ↓    ⤷Sect. Eustigmatopteris C. Chr., 1913
              ↓    ⤷Sect. Fibrillosae Ching, 1938

               ⤷Sous-Genre : Glaphyropteris (Fée) C. Chr., 1911

               ⤷Sous-Genre : Goniopteris (C. Presl) C. Chr., 1905

              ↓    ⤷Sect. Hirtae C. Chr., 1934

              ↓    ⤷Sect. Hirtipedes Fraser-Jenk., 1986

              ↓    ⤷Sect. Indusiatae S.G. Lu, 1999

               ⤷Sous-Genre : Lastrea (Bory) C. Chr., 1911

               ⤷Sous-Genre : Leptogramma (J. Sm.) C. Chr., 1906

              ↓    ⤷Sect. Lophodium (Newman) C. Chr. ex H. Itô, 1939

              ↓    ⤷Sect. Marginatae Fraser-Jenk., 1986

               ⤷Sous-Genre : Meniscium (Schreb.) C. Chr., 1905

              ↓    ⤷Sect. Nephrocystis H. Itô, 1955

              ↓    ⤷Sect. Nothoperanema (Tagawa) Li Bing Zhang, 2012

               ⤷Sous-Genre : Nothoperanema Tagawa, 1938
              ↓    ⤷Sect. Pallidae Fraser-Jenk., 1986

              ↓    ⤷Sect. Pandae Fraser-Jenk., 1986

               ⤷Sous-Genre : Parapolystichum (Keyserl.) C. Chr., 1920

               ⤷Sous-Genre : Peismapodium Maxon, 1909

              ↓   ⤷Sect. Peltochlaena Fée ex C. Chr., 1913

              ↓   ⤷Sect. Peranema (D. Don) Li Bing Zhang, 2012

               ⤷Sous-Genre : Phegopteris (C. Presl) C. Chr., 1905

               ⤷Sous-Genre : Polystichopsis (J. Sm.) C. Chr., 1934

              ↓   ⤷Sect. Purpurascentes Fraser-Jenk., 1986

               ⤷Sous-Genre : Pycnopteris (T. Moore) Ching, 1938
              ↓   ⤷Sect. Remota Fraser-Jenk., 1986

              ↓   ⤷Sect. Shiehia L.-Y. Kuo & Y.H. Chang, 2016

              ↓   ⤷Sect. Splendentes Fraser-Jenk., 1986

               ⤷Sous-Genre : Stegnogramma (Blume) C. Chr., 1906

               ⤷Sous-Genre : Steiropteris C. Chr., 1911

              ↓    ⤷Sect. Stenolepia (Alderw.) L.-Y. Kuo & Y.H. Chang, 2016

               ⤷Sous-Genre : Stigmatopteris (C. Chr.) C. Chr., 1913

                    ⤷Sect. Subincisae C. Chr., 1934
                    ⤷Sect. Variae Fraser-Jenk., 1986

Dryopteris Adans., 1763
✔Dryopteris sect. Acrophorus (C. Presl) Li Bing Zhang, 2012

✔Dryopteris sect. Acrorumohra (H. Itô) Li Bing Zhang & H. He Fl.,

    2013
✔Dryopteris sect. Aemulae Fraser-Jenk., 1986
? Dryopteris sect. Amplae C. Chr., 1934
✔Dryopteris sect. Caespitosae S.G. Lu, 1990
? Dryopteris ser. Championianae Ching ex S.F. Lu, 1999
? Dryopteris sect. Chrysocomae S.G. Lu, 1990
? Dryopteris subg. Ctenitis C. Chr., 1911
? Dryopteris subg. Cyclosorus (Link) C. Chr., 1905
✔Dryopteris sect. Diacalpe (Blume) Li Bing Zhang, 2012
✔Dryopteris sect. Dryopsis (Holttum & P.J. Edwards) Li Bing Zhang,

    2012
? Dryopteris sect. Dryopteris
✔Dryopteris subg. Dryopteris
✔Dryopteris sect. Erythrovariae H. Itô, 1936
✔Dryopteris subg. Erythrovariae (H. Itô) Fraser-Jenk., 1986
? Dryopteris subg. Erythroviae Kraenzl., 1999

? Dryopteris subg. Eudryopteris C. Chr., 1906
? Dryopteris sect. Eustigmatopteris C. Chr., 1913

✔Dryopteris sect. Fibrillosae Ching, 1938
? Dryopteris subg. Glaphyropteris (Fée) C. Chr., 1911
? Dryopteris subg. Goniopteris (C. Presl) C. Chr., 1905
? Dryopteris sect. Hirtae C. Chr., 1934
✔Dryopteris sect. Hirtipedes Fraser-Jenk., 1986
✔Dryopteris sect. Indusiatae S.G. Lu, 1999
? Dryopteris subg. Lastrea (Bory) C. Chr., 1911
? Dryopteris subg. Leptogramma (J. Sm.) C. Chr., 1906
✔Dryopteris sect. Lophodium (Newman) C. Chr. ex H. Itô, 1939
✔Dryopteris sect. Marginatae Fraser-Jenk., 1986
? Dryopteris subg. Meniscium (Schreb.) C. Chr., 1905
✔Dryopteris sect. Nephrocystis H. Itô, 1955
✔Dryopteris sect. Nothoperanema (Tagawa) Li Bing Zhang, 2012
✔Dryopteris subg. Nothoperanema Tagawa, 1938
✔Dryopteris sect. Pallidae Fraser-Jenk., 1986
✔Dryopteris sect. Pandae Fraser-Jenk.,  1986
? Dryopteris subg. Parapolystichum (Keyserl.) C. Chr., 1920
? Dryopteris subg. Peismapodium Maxon, 1909
? Dryopteris sect. Peltochlaena Fée ex C. Chr., 1913
✔Dryopteris sect. Peranema (D. Don) Li Bing Zhang, 2012
? Dryopteris subg. Phegopteris (C. Presl) C. Chr., 1905
? Dryopteris subg. Polystichopsis (J. Sm.) C. Chr., 1934
✔Dryopteris sect. Purpurascentes Fraser-Jenk., 1986
✔Dryopteris subg. Pycnopteris (T. Moore) Ching, 1938
✔Dryopteris sect. Remota Fraser-Jenk., 1986
? Dryopteris sect. Shiehia L.-Y. Kuo & Y.H. Chang, 2016
✔Dryopteris sect. Splendentes Fraser-Jenk., 1986
? Dryopteris subg. Stegnogramma (Blume) C. Chr., 1906
? Dryopteris subg. Steiropteris C. Chr., 1911
? Dryopteris sect. Stenolepia (Alderw.) L.-Y. Kuo & Y.H. Chang, 2016
? Dryopteris subg. Stigmatopteris (C. Chr.) C. Chr., 1913
? Dryopteris sect. Subincisae C. Chr., 1934
✔Dryopteris sect. Variae Fraser-Jenk., 1986

✔= légitime (accepté).

 

Espèces :

En orange, les espèces légitimes répertoriées

et enregistrées chez Kew.

Source :

 

Pour certains,

1964 espèces enregistrées sous le nom

de genre Dryopteris Adans.,

plus 28 hybrides

196 espèces légitimes (acceptées),

plus 1 hybride pour certains,

peut être plus/moins d'espèces pour d'autres.

Dryopteris acrophorus Li Bing Zhang, 2012

Dryopteris acuminatum Watts,   1916
Dryopteris acutodentata Ching, 1938

Dryopteris adscendens (Ching) Li Bing Zhang, 2012

Dryopteris affinis (Lowe) Fraser-Jenk., 1979
Dryopteris africana (Desv.) C. Chr., 1905

Dryopteris alpestris Tagawa, 1934

Dryopteris alpicola Ching & Z.R. Wang, 1985
Dryopteris amurensis Christ, 1909

Dryopteris angustifrons (Hook.) Kuntze, 1891

Dryopteris apiciflora (Wall. ex Mett.) Kuntze, 1891
Dryopteris arguta (Kaulf.) Watt, 1866
Dryopteris assamensis (C. Hope) C. Chr. & Ching, 1933
Dryopteris asterothrix Rosenst., 1909
Dryopteris atrata (Wall. ex Kunze) Ching, 1933

Dryopteris barbigera (T. Moore ex Hook.) Kuntze, 1891
Dryopteris blanfordii (C. Hope) C. Chr., 1905
Dryopteris bodinieri (Christ) C. Chr., 1905

Dryopteris cacaina Tagawa, 1937
Dryopteris canelensis Rosenst., 1909

Dryopteris caroli-hopei Fraser-Jenk., 1989
Dryopteris carthusiana (Vill.) H.P. Fuchs, 1959
Dryopteris championii (Benth.) C. Chr. ex Ching, 1933
Dryopteris chinensis (Baker) Koidz., 1930
Dryopteris christensenae (Ching) Li Bing Zhang, 2012
Dryopteris christii C. Chr., 1905
Dryopteris chrysocoma (Christ) C. Chr., 1905
Dryopteris clarkei (Baker) Kuntze, 1891
Dryopteris cochleata (Buch.-Ham. ex D. Don) C. Chr., 1905

Dryopteris commixta Tagawa, 1933

Dryopteris conjugata Ching, 1941
Dryopteris coreano-montana Nakai, 1921
Dryopteris costalisora Tagawa, 1934
Dryopteris crassirhizoma Nakai, 1920
Dryopteris cycadina (Franch. & Sav.) C. Chr., 1905
Dryopteris cyclopeltidiformis C. Chr., 1937

Dryopteris decipiens (Hook.) Kuntze, 1891
Dryopteris dehuaensis Ching & K.H. Shing, 1991
Dryopteris diacalpe Li Bing Zhang, 2012
Dryopteris dickinsii (Franch. & Sav.) C. Chr., 1905

Dryopteris diffracta (Baker) C. Chr., 1905
Dryopteris dilatata (Hoffm.) A. Gray, 1848
Dryopteris diplazioides Kuntze, Revis. Gen. Pl. 2: 812 1891
Dryopteris dulongensis (S.K. Wu & X. Cheng) Li Bing Zhang, 2012

Dryopteris emeiensis (Ching) Li Bing Zhang, 2012
Dryopteris enneaphylla (Baker) C. Chr., 1905
Dryopteris erythrosora (D.C. Eaton) Kuntze, 1891
Dryopteris expansa (C. Presl) Fraser-Jenk. & Jermy, 1977
Dryopteris exstipellata (Ching & S.H. Wu) Li Bing Zhang, 2012

Dryopteris falcata (L. f.) Kuntze, 1891

Dryopteris feei C. Chr., 1905
Dryopteris fibrillosa (Baker) Christ, 1905
Dryopteris fibrillosissima Ching Fl. Xizang., 1983
Dryopteris filix-mas (L.) Schott, 1834
Dryopteris formosana (Christ) C. Chr., 1905
Dryopteris fructuosa (Christ) C. Chr., 1905
Dryopteris fuscipes C. Chr., 1917

Dryopteris gemmifera S.Y. Dong, 2010
Dryopteris gigantea (Blume) Kuntze, 1891
Dryopteris goeringiana (Kunze) Koidz., 1929

Dryopteris grandifrons Li Bing Zhang, 2012
Dryopteris guadalupensis (Fée) Kuntze, 1891
Dryopteris guangxiensis S.G. Lu, 1999
Dryopteris gymnophylla (Baker) C. Chr., 1905
Dryopteris gymnosora (Makino) C. Chr., 1905

Dryopteris habaensis Ching, 1941
Dryopteris handeliana C. Chr., 1937
Dryopteris hangchowensis Ching, 1938
Dryopteris hasseltii (Blume) C. Chr., 1905
Dryopteris hendersonii (Bedd.) C. Chr., 1905

Dryopteris heterolaena C. Chr., 1924
Dryopteris himachalensis Fraser-Jenk., 1989
Dryopteris hirsutosetosa Hieron., 1907
Dryopteris holttumii Li Bing Zhang, 2013
Dryopteris hondoensis Koidz., 1932
Dryopteris hookeriana (T. Moore) Li Bing Zhang, 2012

Dryopteris immixta Ching, 1974

Dryopteris indusiata (Makino) Makino & Yamam., 1932
Dryopteris integriloba C. Chr., 1933

Dryopteris juergensii (Rosenst.) C. Chr., 1906

Dryopteris juxtaposita Christ, 1907

Dryopteris kawakamii Hayata, 1911
Dryopteris kinkiensis Koidz. ex Tagawa, 1933
Dryopteris komarovii Kossinsky, 1921
Dryopteris kungiana Li Bing Zhang, 2012
Dryopteris kwanzanensis Tagawa, 1938

Dryopteris labordei (Christ) C. Chr., 1906
Dryopteris lacera (Thunb.) Kuntze, 1891
Dryopteris lachoongensis (Bedd.) B.K. Nayar & S. Kaur, 1972
Dryopteris latibasis Ching, 1941

Dryopteris lepidopoda Hayata, 1914
Dryopteris lepidorachis C. Chr., 1906
Dryopteris liangkwangensis Ching, 1937
Dryopteris liboensis P.S. Wang, X.Y. Wang & Li Bing Zhang,  2012
Dryopteris lunanensis (Christ) C. Chr., 1905

Dryopteris marginata (C.B. Clarke) Christ, 1907
Dryopteris maximowicziana (Miq.) C. Chr., 1924
Dryopteris medogensis (Ching & S.K. Wu) Li Bing Zhang, 2012
Dryopteris melanocarpa Hayata, 1914
Dryopteris meniscioides (Willd.) Kuntze, 1891

Dryopteris microlepis (Baker) C. Chr., 1913
Dryopteris mollis (Sw.) Hieron., 1907
Dryopteris monticola (Makino) C. Chr., 1905
Dryopteris montigena Ching, 1941

Dryopteris namegatae (Sa. Kurata) Sa. Kurata, 1969
Dryopteris neorosthorni Ching, 1941
Dryopteris nobilis Ching, 1941
Dryopteris nodosa (C. Presl) Li Bing Zhang,  2012
Dryopteris nyingchiensis Ching, 1983

Dryopteris paleacea (T. Moore) Hand.-Mazz., 1908

Dryopteris paleolata (Pic. Serm.) Li Bing Zhang, 2012
Dryopteris panda (C.B. Clarke) Christ, 1909
Dryopteris papilio (C. Hope) C. Chr., 1905
Dryopteris parafuscipes Ching & Z.Y. Liu, 1984
Dryopteris paralunanensis W.M. Chu ex S.G. Lu, 1991
Dryopteris patens (Sw.) Kuntze, 1891
Dryopteris pellucidopunctata C. Chr., 1905
Dryopteris peninsulae Kitag., 1935
Dryopteris peranema Li Bing Zhang, 2012
Dryopteris phegopteris (L.) C. Chr., 1905

Dryopteris pilosohispida (Hook.) C. Chr., 1913
Dryopteris pilosula (Klotzsch & H. Karst. ex Mett.) Hieron., 1907
Dryopteris podophylla (Hook.) Kuntze, 1891
Dryopteris polita Rosenst., 1914
Dryopteris porosa Ching, 1938
Dryopteris porphyricola Copel., 1912
Dryopteris pseudocaenopteris (Kunze) Li Bing Zhang, 2012
Dryopteris pseudolunanensis Tagawa, 1940
Dryopteris pseudosparsa Ching, 1938
Dryopteris pteridiiformis Christ, 1907

Dryopteris pulcherrima Ching, 1938
Dryopteris pulvinulifera (Bedd.) Kuntze, 1891
Dryopteris pycnopteroides (Christ) C. Chr., 1913

Dryopteris redactopinnata S.K. Basu & Panigrahi, 1980
Dryopteris reflexosquamata Hayata, 1914
Dryopteris remota (A. Braun) Hayek, 1908
Dryopteris remotipinnulata Bonap., 1925
Dryopteris robertiana (Hoffm.) C. Chr., 1905
Dryopteris rosthornii (Diels) C. Chr., 1905
Dryopteris rubrobrunnea W.M. Chu, 1992

Dryopteris ryo-itoana Sa. Kurata, 1967

Dryopteris sacrosancta Koidz., 1924
Dryopteris saxifraga H. Itô, 1936
Dryopteris scottii (Bedd.) Ching, 1933
Dryopteris sericea C. Chr., 1913
Dryopteris serratodentata (Bedd.) Hayata, 1914
Dryopteris setigera (Blume) Kuntze, 1891
Dryopteris setosa (Thunb.) Akas., 1959
Dryopteris shikokiana (Makino) C. Chr., 1905
Dryopteris sieboldii (Van Houtte ex Mett.) Kuntze, 1891

Dryopteris sikkimensis (Bedd.) Kuntze, 1891
Dryopteris simasakii (H. Itô) Sa. Kurata, 1970
Dryopteris sinofibrillosa Ching, 1940
Dryopteris sordidipes Tagawa, 1934
Dryopteris sparsa (D. Don) Kuntze, 1891
Dryopteris sphaeropteroides (Baker) C. Chr., 1905
Dryopteris splendens (Hook.) Kuntze, 1891
Dryopteris squamifera Ching & S.K. Wu Fl. Xizang., 1983
Dryopteris squamiseta (Hook.) Kuntze, 1891
Dryopteris stenolepis (Baker) C. Chr., 1905

Dryopteris stenophylla (Sodiro) C. Chr., 1905
Dryopteris subatrata Tagawa, 1940
Dryopteris subexaltata (Christ) C. Chr., 1905
Dryopteris subimpressa Loyal, 1969
Dryopteris sublacera Christ, 1909
Dryopteris submarginata Rosenst., 1914
Dryopteris subpycnopteroides Ching ex Fraser-Jenk.,1986
Dryopteris subreflexipinna Ogata, 1935
Dryopteris subtriangularis (C. Hope) C. Chr., 1906
Dryopteris subtripinnata (Miq.) Kuntze, 1891

Dryopteris tahmingensis Ching, 1938
Dryopteris tenuicula C.G. Matthew & Christ, 1909

Dryopteris tenuipes (Rosenst.) Seriz., 1971
Dryopteris tetragona (Sw.) Kuntze, 1891
Dryopteris thibetica (Franch.) C. Chr., 1905
Dryopteris tingiensis Ching & S.K. Wu ex Fraser-Jenk., 1986
Dryopteris tokyoensis (Matsum. ex Makino) C. Chr., 1905
Dryopteris toyamae Tagawa, 1939
Dryopteris transmorrisonensis (Hayata) Hayata, 1914
Dryopteris tsoongii Ching, 1987

Dryopteris × uliginosa (A. Braun ex Döll) Kuntze ex Druce, 1909
Dryopteris uniformis (Makino) Makino, 1909
Dryopteris unita (L.) Kuntze, 1891

Dryopteris varia (L.) Kuntze, 1891

Dryopteris wallichiana (Spreng.) Hyl., 1953
Dryopteris whangshangensis Ching, 1938
Dryopteris woodsiisora Hayata, 1916
Dryopteris wusugongii Li Bing Zhang, 2012
Dryopteris wuyishanica Ching & P.C. Chiu, 1987
Dryopteris wuzhaohongii Li Bing Zhang, 2012

Dryopteris xholttumii Li Bing Zhang, 2013
Dryopteris xunwuensis Ching & K.H. Shing, 1990

Dryopteris yigongensis Ching 1983
Dryopteris yongdeensis W.M. Chu ex S.G. Lu, 1990
Dryopteris yoroii Seriz., 1971
Dryopteris yungtzeensis Ching, 1941

Dryopteris zhuweimingii Li Bing Zhang, 2012

Les Hybrides :

Dryopteris × uliginosa

Source :

344 espèces

Le genre Dryopteris 3/4

De İ à O

İ/

Dryopteris immixta Ching, 1974
Dryopteris inaequalis (Schltdl.) Kuntze, 1891
Dryopteris indonesiana Darnaedi, M.Kato & K.Iwats., 1989
Dryopteris indusiata Makino & Yamam., 1932
Dryopteris × initialis Fraser-Jenk. & Corley, 1972

    (D. caucasica × D. oreades)

Dryopteris insularis Kodama, 1913
Dryopteris integriloba C.Chr., 1933
Dryopteris integripinnula Ching, 1981
Dryopteris intermedia A.Gray, 1848

⤷ intermedia subsp. azorica (Christ) Jermy, 1984
⤷ intermedia subsp. maderensis (Alston) Fraser-Jenk., 1999

J/

Dryopteris jinpingensis Z.Y.Zuo, J.Mei Lu & D.Z.Li, 2024
Dryopteris jishouensis G.X.Chen & D.G.Zhang, 2013
Dryopteris jiucaipingensis P.S.Wang, Q.Luo & Li Bing Zhang, 2012
Dryopteris juxtaposita Christ, 1907

K/

Dryopteris karwinskyana Kuntze, 1891
Dryopteris kashmiriana Fraser-Jenk. & Widén, 2015
Dryopteris katangaensis J.P.Roux, 2003
Dryopteris kawakamii Hayata, 1911
Dryopteris khullarii Fraser-Jenk., 1989
Dryopteris kilemensis (Kuhn) Kuntze, 1891
Dryopteris kinkiensis Koidz. ex Tagawa, 1933
Dryopteris kinokuniensis Sa.Kurata, 1970
Dryopteris knoblochii A.R.Sm., 2004
Dryopteris kobayashii Kitag., 1935
Dryopteris koidzumiana Tagawa, 1933
Dryopteris komarovii Kossinsky, 1921
Dryopteris × kominatoensis Tagawa, 1937

    (D. monticola × D. tokyoensis)

Dryopteris × kouzaii Akasawa, 1979

    (D. dickinsii × D. handeliana)

Dryopteris kuratae Nakaike ex Hoshiz. & K.A.Wilson, 1999
Dryopteris kwanzanensis Tagawa, 1938

L/
Dryopteris labordei (Christ) C.Chr., 1905
Dryopteris lacera (Thunb.) Kuntze, 1891
Dryopteris laeta (Kom.) C.Chr., 1905
Dryopteris × lawalreei Janch., 1963

    (D. borreri × D. carthusiana × D. filix-mas)

Dryopteris × leedsii Wherry, 1942

    (D. celsa × D. marginalis)

Dryopteris lepidopoda Hayata, 1914
Dryopteris lepidorachis C.Chr., 1905
Dryopteris lepinei (Kuhn) Kuntze, 1891
Dryopteris leucorhachis (Cheeseman) C.Chr., 1905
Dryopteris lewalleana Pic.Serm., 1985
Dryopteris liankwangensis Ching, 1937
Dryopteris liboensis P.S.Wang, X.Y.Wang & Li Bing Zhang, 2012
Dryopteris × liddarensis Fraser-Jenk., 1989

    (D. barbigera × D. serratodentata)

Dryopteris × ligustica Gibby, S.Jess. & Marchetti, 2024

    (D. filix-mas × D. tyrrhena)

Dryopteris lijianxiui Xiao J.Li, 2021
Dryopteris × loyalii Fraser-Jenk., 1989

    (D. caroli-hopei × D. marginata)

Dryopteris ludoviciana (Kunze) Small, 1938
Dryopteris lunanensis (Christ) C.Chr., 1905
Dryopteris lustrata C.Chr., 1905

M/
Dryopteris × macdonellii Fraser-Jenk., 1989

    (D. blanfordii × D. filix-mas)

Dryopteris macrochlamys (Fée) Fraser-Jenk., 1989
Dryopteris macrolepidota Copel., 1932
Dryopteris macropholis Lorence & W.L.Wagner, 2011
Dryopteris × madalenae Fraser-Jenk., 1981

    (D. crispifolia × D. intermedia subsp. azorica)

Dryopteris × makabensis Kaor.Yamam. & N.Murak., 2024

    (D. caudipinna × D. tokyoensis)

Dryopteris mangindranensis Tardieu, 1956
Dryopteris manipurensis (Bedd.) C.Chr., 1905
Dryopteris manniana (Hook.) C.Chr., 1905
Dryopteris × mantoniae Fraser-Jenk. & Corley, 1972

    (D. filix-mas × D. oreades)

Dryopteris marginalis (L.) A.Gray, 1848
Dryopteris marginata (C.B.Clarke) Christ, 1907
Dryopteris × martinsiae Fraser-Jenk., 1981

    (D. aemula × D. crispifolia)

Dryopteris mauiensis C.Chr., 1905
Dryopteris maximowicziana (Miq.) C.Chr., 1924
Dryopteris maximowiczii (Baker) Kuntze, 1891
Dryopteris maxonii Underw. & C.Chr., 1911
Dryopteris × mayebarae Tagawa, 1938

    (D. commixta × D. uniformis)

Dryopteris medioxima Koidz., 1932
Dryopteris meghalaica Fraser-Jenk. & Gibby, 1999
Dryopteris melanocarpa Hayata, 1914
Dryopteris mengshanensis J.X.Li & Xiao J.Li, 2024
Dryopteris × mickelii J.H.Peck, 2001

    (D. clintoniana × D. goldieana)

Dryopteris microlepis (Baker) C.Chr., 1913
Dryopteris mindshelkensis Pavlov, 1954
Dryopteris × mituii Seriz., 1971

    (D. lacera × D. uniformis)

Dryopteris × miyazakiensis Miyam., 1984

    (D. commixta × D. hangchowensis)

Dryopteris × montgomeryi Fraser-Jenk. & Widén, 2006

    (D. filix-mas × D. marginalis)

Dryopteris monticola (Makino) C.Chr., 1905
Dryopteris montigena Ching, 1941
Dryopteris munchii A.R.Sm., 1975

N/

Dryopteris namegatae (Sa.Kurata) Sa.Kurata, 1969
Dryopteris napoleonis (Bory) Kuntze, 1891
Dryopteris neoassamensis Ching, 1938
Dryopteris × neowherryi W.H.Wagner, 1966

    (D. goldieana × D. marginalis)

Dryopteris nidus (Baker) Li Bing Zhang, 2012
Dryopteris nigropaleacea (Fraser-Jenk.) Fraser-Jenk., 1982
Dryopteris nipponensis Koidz., 1931
Dryopteris nodosa (C.Presl) Li Bing Zhang, 2012
Dryopteris nubigena Maxon & C.V.Morton, 1937

O/
Dryopteris obtusiloba (Baker) Kuntze, 1891
Dryopteris obtusipinnula (Fraser-Jenk.) Fraser-Jenk., 2023
Dryopteris occidentalis J.P.Roux, 2004
Dryopteris odontoloma (T.Moore ex Bedd.) C.Chr., 1924
Dryopteris okinawensis H.Itô, 1935
Dryopteris oligodonta (Desv.) Pic.Serm., 1951
Dryopteris oreades Fomin, 1910 publ. 1911

⤷ oreades subsp. corsica Fraser-Jenk., 2024

Dryopteris × orexpansa Pérez Carro & Fern.Areces, 2013

    (D. expansa × D. oreades)

Dryopteris × otomasui Sa.Kurata, 1967
    (D. commixta × D. polylepis)




⏪⏩

Le genre Dryopteris 2/4

Source :

344 espèces enregistrées sous le nom

de genre Dryopteris Adans.,

344 espèces légitimes (acceptées),

plus 88 hybrides plus 29 sous-espèces,

18 variétés, 6 formes.

Dryopteris se décline également par des hybridations

entre sous-espèce, variété ou forme.

Ces hybridations plus complexes sont désignés par :

-  6 nothosubsp. ( hybride entre ou avec au moins 1 sous-espèce),

- 5 nothovar. (hybride entre ou  avec au moins 1 variété)

- 1 nothof. (hybride entre ou avec au moins 1 forme)

De A à H

A/
Dryopteris acutissima H.J.Wei & Z.Y.Zuo, 2025

Dryopteris acutodentata Ching, 1938
Dryopteris adnata (Blume) Alderw., 1909
Dryopteris aemula (Aiton) Kuntze, 1891

Dryopteris affinis (Lowe) Fraser-Jenk., 1979, nom. cons.

⤷ affinis subsp. atropes Fraser-Jenk., 2022
⤷ affinis subsp. borreri (Newman) Fraser-Jenk., 1980
⤷ affinis subsp. cambrensis Fraser-Jenk., 1987
⤷ affinis subsp. jessenii (Fraser-Jenk.) Fraser-Jenk., 2007
⤷ affinis subsp. persica Fraser-Jenk., 1980
⤷ affinis subsp. pontica Fraser-Jenk., 1996
⤷ affinis subsp. pseudocomplexa (Fraser-Jenk.) Fraser-Jenk., 2022
⤷ affinis subsp. pseudodisjuncta (Fraser-Jenk.) Fraser-Jenk., 1996
 ⤷ affinis var. austrogallica Fraser-Jenk. & Boudrie, 2024
 ⤷ affinis var. distans (Viv.) Fraser-Jenk., 2024
 ⤷ affinis var. kerryensis Fraser-Jenk., 1996
 ⤷ affinis var. paleaceocrispa (T.Moore) Fraser-Jenk., 2021
 ⤷ affinis var. paleaceolobata (T.Moore) Fraser-Jenk., 2021
 ⤷ affinis var. splendens (Ehrler ex Becherer) Fraser-Jenk., 1980

 ⤷ affinis f. azorica (Fraser-Jenk.) Fraser-Jenk., 2024
 ⤷ affinis f. cluthensis (A.R.Church) Fraser-Jenk., 2024
 ⤷ affinis f. dilatata (Luerss.) Fraser-Jenk., 2024)
 ⤷ affinis f. picoensis Fraser-Jenk., 2024

Dryopteris aitoniana Pic.Serm., 1951
Dryopteris × alejandrei Pérez Carro & Fern.Areces, 2007

   (D. mindshelkensis × D. oreades)

Dryopteris × algonquinensis D.M.Britton, 1975

    (D. fragrans × D. marginalis)

Dryopteris alpestris Tagawa, 1934
Dryopteris × alpirsbachensis Freigang, Zenner, Bujnoch,

  S.Jess. & Magauer, 2017  (D. carthusiana × D. remota)

Dryopteris amblyodonta J.P.Roux, 2004
Dryopteris × ambroseae Fraser-Jenk. & Jermy, 1977

    (D. dilatata × D. expansa)

Dryopteris amurensis (Milde ) Christ, 1909
Dryopteris aneitensis (Hook.) C.Chr., 1905
Dryopteris angustifrons (T.Moore) Kuntze, 1891
Dryopteris angustipalea Darnaedi, M.Kato & K.Iwats., 1989
Dryopteris annamensis (Tagawa) Li Bing Zhang, 2012
Dryopteris antarctica (Baker) C.Chr., 1934
Dryopteris anthracinisquama Miyam., 1984
Dryopteris apiciflora (Wall. ex Mett.) Kuntze, 1891
Dryopteris approximata Sledge, 1973
Dryopteris × apuana Gibby, S.Jess. & Marchetti, 1996

    (D. mindshelkensis × D. villarii)

Dryopteris aquilinoides (Desv.) C.Chr., 1905
Dryopteris arborescens (Baker) Kuntze, 1891
Dryopteris ardechensis Fraser-Jenk., 1981
Dryopteris × arecesiae Pérez Carro & T.E.Díaz, 1990

    (D. aemula × D. corleyi)

Dryopteris arguta (Kaulf.) Watt, 1867
Dryopteris arunachalensis (Fraser-Jenk. & Benniamin)

  R.Kr.Singh & V.K.Rawat, 2024
Dryopteris ascensionis (Hook.) Kuntze, 1891
Dryopteris assamensis (C.Hope) C.Chr. & Ching, 1933
Dryopteris × asturiensis Fraser-Jenk. & Gibby, 1986

    (D. corleyi × D. pseudomas)

Dryopteris athamantica (Kunze) Kuntze, 1891
Dryopteris atrata (Wall. ex Kunze) Ching, 1933
Dryopteris aurantiaca J.P.Roux, 2005
Dryopteris × australis (Wherry) Small, 1938

    (D. celsa × D. ludoviciana)

Dryopteris austroindica Fraser-Jenk., 1989

B/

Dryopteris bamleriana Rosenst., 1912

Dryopteris baniensis Rosenst., 1909
Dryopteris barbigera (Hook.) Kuntze, 1891
Dryopteris basisora Christ, 1909
Dryopteris × benedictii (Farw.) Wherry, 1961

    (D. carthusiana × D. clintoniana)

Dryopteris bernieri Tardieu, 1956
Dryopteris bhutanica (Fraser-Jenk. & Pariyar) Fraser-Jenk., 2023
Dryopteris bissetiana (Baker) C.Chr., 1905
Dryopteris blanfordii (C.Hope) C.Chr., 1905

⤷ blanfordii subsp. nigrosquamosa (Ching) Fraser-Jenk., 1989

Dryopteris bodinieri (Christ) C.Chr., 1905
Dryopteris bojeri Kuntze, 1891
Dryopteris × boottii (Tuck.) Underw., 1893

    (D. cristata × D. intermedia)

Dryopteris borbasii Litard., 1910
Dryopteris × brathaica Fraser-Jenk. & Reichst., 1977

    (D. carthusiana × D. filix-mas)

Dryopteris × burgessii B.Boivin, 1966

    (D. clintoniana × D. marginalis)

Dryopteris burnatii Christ & Wilczek, 1913

C/

Dryopteris cacaina Tagawa, 1937
Dryopteris campyloptera Clarkson, 1930
Dryopteris camusiae Fraser-Jenk., 1997
Dryopteris × cantabrica Alejandre, Pérez Carro & Fern.Areces,

  2016          (D. corleyi × D. guanchica)
Dryopteris caperata J.P.Roux, 2005

Dryopteris caroli-hopei Fraser-Jenk., 1989
Dryopteris carpatica S.Jess. & Bujnoch, 2022
Dryopteris carthusiana (Vill.) H.P.Fuchs, 1959

 ⤷ carthusiana var. enigmatica Harmaja, 2012

Dryopteris caucasica (A.Braun) Fraser-Jenk. & Corley, 1972
Dryopteris caudipinna Nakai, 1931
Dryopteris × cedroensis Gibby & Widén, 1983

    (D. corleyi × D. guanchica)

Dryopteris celsa (W.Palmer) Knowlt., T.S.Palmer & Pollard ex Small,

  1900
Dryopteris chaerophyllifolia (Zippel) C.Chr., 1905
Dryopteris championii (Benth.) C.Chr., 1933
Dryopteris chichisimensis Nakai ex H.Itô, 1935
Dryopteris chinensis (Baker) Koidz., 1930
Dryopteris christenseniae (Ching) Li Bing Zhang, 2012
Dryopteris chrysocoma (Christ) C.Chr., 1905
Dryopteris cicatricata J.P.Roux, 2005
Dryopteris cinnamomea (Cav.) C.Chr., 1911
Dryopteris clarkei (Baker) Kuntze, 1891
Dryopteris clintoniana (D.C.Eaton) Dowell, 1906
Dryopteris cochleata (D.Don) C.Chr., 1905
Dryopteris cognata (C.Presl) Kuntze, 1891
Dryopteris collucata Charit., 2022
Dryopteris commixta Tagawa, 1933
Dryopteris comorensis (Tardieu) Fraser-Jenk., 1994
Dryopteris × complexa Fraser-Jenk., 1987

    (D. affinis × D. filix-mas)

⤷ × complexa nothosubsp. complanata (Fraser-Jenk.) Fraser-Jenk.,
   2024      (D. affinis subsp. pseudodisjuncta × D. filix-mas)

⤷ × complexa nothosubsp. convoluta (Fraser-Jenk.) Fraser-Jenk.,

   2024       (D. affinis subsp. cambrensis × D. filix-mas)

⤷ × complexa nothosubsp. critica Fraser-Jenk., 1987

   (D. affinis subsp. borreri × D. filix-mas)
⤷ × complexa nothosubsp. inconspicua (Fraser-Jenk.) Fraser-Jenk.,

   2022        (D. affinis subsp. pseudocomplexa × D. filix-mas)
⤷ × complexa nothosubsp. transsilvanica Tlałka, S.Jess., A.Rostański

   & Rojek, 2019       (D. affinis subsp. jessenii × D. filix-mas)

⤷ × complexa nothovar. contorta (Fraser-Jenk.) Fraser-Jenk., 2024

   (D. affinis var. paleaceolobata × D. filix-mas)

⤷ × complexa nothovar. eschelmuelleri (Fraser-Jenk.) Fraser-Jenk. &

   Parris, 2020 pub. 2021   (D. affinis var. splendens × D. filix-mas)

⤷ × complexa nothovar. hibernica Fraser-Jenk. & Trewren, 2024

   (D. affinis var. kerryensis × D. filix-mas)

⤷ × complexa nothovar. occidentalis (Fraser-Jenk.) Fraser-Jenk., 2024
   (D. affinis var. paleaceocrispa × D. filix-mas)
⤷ × complexa nothovar. rufosquama Fraser-Jenk., 2024
   (D. affinis var. insubrica × D. filix-mas)

⤷ × complexa nothof. urbachensis (S.Jess. & Bujnoch) Fraser-Jenk.,
   2024    (D. affinis f. dilatata × D. filix-mas)

Dryopteris conjugata Ching, 1941
Dryopteris conversa Alderw., 1915
Dryopteris coreanomontana Nakai, 1921
Dryopteris corleyi Fraser-Jenk., 
Dryopteris × correllii W.H.Wagner & A.V.Gilman, 2001

    (D. carthusiana × D. goldieana)

Dryopteris costalisora Tagawa, 1934
Dryopteris crassirhizoma Nakai, 1920
Dryopteris crinalis (Hook. & Arn.) C.Chr., 1905

 ⤷ crinalis var. podosora (W.H.Wagner) D.D.Palmer, 2002

Dryopteris crispifolia Rasbach, Reichst. & G.Vida, 1977
Dryopteris cristata (L.) A.Gray, 1848
Dryopteris cycadina (Franch. & Sav.) C.Chr., 1905
Dryopteris cyclopeltidiformis C.Chr., 1937
Dryopteris cyclosorus Alderw., 1924

D/

Dryopteris damingshanensis Li Bing Zhang & H.M.Liu, 2014
Dryopteris daozhenensis P.S.Wang & X.Y.Wang, 2001
Dryopteris decipiens (Hook.) Kuntze, 1891
Dryopteris dehuaensis Ching & K.H.Shing, 1982
Dryopteris deparioides (T.Moore) Kuntze, 1891

⤷ deparioides subsp. ambigua (Sledge) Fraser-Jenk., 1989
⤷ deparioides subsp. concinna C.Chr., 1934

Dryopteris devriesei Rosenst., 1917
Dryopteris × deweveri (Jansen) Jansen & Wachter, 1934

    (D. carthusiana × D. dilatata)

Dryopteris diacalpioides (Ching) Li Bing Zhang, 2012
Dryopteris dibangensis C.Chanda & Fraser-Jenk., 2024

Dryopteris dickinsii (Franch. & Sav.) C.Chr., 1905
Dryopteris dicksonioides (Mett.) Copel., 1932
Dryopteris diffracta (Baker) C.Chr., 1905
Dryopteris dilatata (Hoffm.) A.Gray, 1848

⤷ dilatata subsp. lusitanica Fraser-Jenk., Andreis & Vasco Silva, 2021

Dryopteris × doeppii Rothm., 1945

    (D. affinis × A. carthusiana)

Dryopteris × doluchanovii Askerov, 1979

    (D. dilatata × D. pseudomas)

Dryopteris × dowellii (Farw.) Wherry, 1961

    (D. clintoniana × D. intermedia)

Dryopteris dracomontana Schelpe & N.C.Anthony, 1982
Dryopteris drakei C.Chr., 1905
Dryopteris dulongensis (S.K.Wu & X.Cheng) Li Bing Zhang, 2012

E/
Dryopteris edwardsii Fraser-Jenk., 1997
Dryopteris enneaphylla (Baker) C.Chr., 1905
Dryopteris erythrosora (D.C.Eaton) Kuntze, 1891
Dryopteris erythrovaria K.Hori & N.Murak., 2018
Dryopteris × euxinensis Fraser-Jenk. & Corley, 1972

    (D. caucasica × D. filix-mas)

Dryopteris expansa (C.Presl) Fraser-Jenk. & Jermy, 1977

 ⤷ expansa var. alpina (T.Moore) Viane, 1986
 ⤷ expansa var. willeana (Lid) Elven, 1996

F/
Dryopteris fadenii Pic.Serm., 1984
Dryopteris fangii Ching, Fraser-Jenk. & Z.R.Wang, 1985
Dryopteris fengiana (Ching) ined.
Dryopteris ferruginea (Baker) Kuntze, 1891
Dryopteris filipaleata J.P.Roux, 2004
Dryopteris filix-mas (L.) Schott, 1834

⤷ filix-mas subsp. brittonii Fraser-Jenk. & Widén, 2006
⤷ filix-mas subsp. pseudorigida (Christ) Askerov, 1983

Dryopteris flaccisquama A.Rojas, 2001
Dryopteris flemingii Fraser-Jenk., 1989
Dryopteris formosana (Christ) C.Chr., 1905
Dryopteris fragrans (L.) Schott, 1834
Dryopteris fragrantiformis Tzvelev, 1989
Dryopteris × fraser-jenkinsii Gibby & Widén, 1983

    (D. affinis × D. guanchica

Dryopteris fructuosa (Christ) C.Chr., 1905
Dryopteris × fujipedis Sa.Kurata, 1965

    (D. crassirhizoma × D. lacera)

Dryopteris × furadensis Bennert, Rasbach, K.Rasbach & Viane, 1991

    (D. aitoniana × D. intermedia subsp. maderensis)

Dryopteris fuscipes C.Chr., 1917

Dryopteris fuscoatra (Hillebr.) W.J.Rob., 1912

 ⤷ fuscoatra var. lamoureuxii Fraser-Jenk., 1994

Dryopteris futura A.R.Sm., 1975

G/
Dryopteris gamblei (C.Hope) C.Chr., 1905
Dryopteris gaoligongensis Z.Y.Zuo, J.Mei Lu & D.Z.Li,

  2021 publ. 2022 
Dryopteris gemmifera S.Y.Dong, 2010
Dryopteris × ghatakii Fraser-Jenk., 1989

    (D. cochleata × D. sparsa subsp. rectipinnula)

Dryopteris glabra (Brack.) Kuntze, 1891

 ⤷ glabra var. alboviridis (W.H.Wagner) D.D.Palmer, 2002
 ⤷ glabra var. flynnii D.D.Palmer, 2002
 ⤷ glabra var. hobdyana (W.H.Wagner) D.D.Palmer, 2002

 ⤷ glabra var. nuda (Underw. ex A.Heller) Fraser-Jenk., 1994
 ⤷ glabra var. pusilla (Hillebr.) Fraser-Jenk., 1994

 ⤷ glabra var. soripes (Hillebr.) Herat ex Fraser-Jenk., 1994

Dryopteris glandulosopaleata J.P.Roux, 2004
Dryopteris goeringiana (Kunze) Koidz., 1929
Dryopteris goldieana (Hook. ex Goldie) A.Gray, 1848
Dryopteris × gomerica Gibby & Widén, 1983

    (D. aemula × D. guanchica)

Dryopteris gonggaensis H.S.Kung, Li Bing Zhang & X.S.Guo, 1995
Dryopteris gorgonea J.P.Roux, 2004
Dryopteris × gotenbaensis Nakaike, 1969

    (D. hondoensis × D. uniformis)

Dryopteris × graeca Fraser-Jenk. & Gibby, 1980

    (D. mindshelkensis × D. pallida)

Dryopteris grandifrons Li Bing Zhang, 2012
Dryopteris guanchica Gibby & Jermy, 1977
Dryopteris guangxiensis S.G.Lu, 1999
Dryopteris gymnophylla (Baker) C.Chr., 1905
Dryopteris gymnosora (Makino) C.Chr., 1905

H/
Dryopteris habaensis Ching, 1941
Dryopteris hadanoi Sa.Kurata, 1966
Dryopteris × haganecola Sa.Kurata, 1958

    (D. cycadina × D. uniformis)

Dryopteris × hakonecola Sa.Kurata, 1958

    (D. dickinsii × D. uniformis)

Dryopteris handeliana C.Chr., 1937
Dryopteris hangchowensis Ching, 1938
Dryopteris hasseltii (Blume) C.Chr., 1905
Dryopteris hawaiiensis (Hillebr.) W.J.Rob., 1912
Dryopteris hayatae Tagawa, 1932
Dryopteris hendersonii (Bedd.) C.Chr., 1905
Dryopteris herbacea Alderw., 1915
Dryopteris heterolaena C.Chr., 1924
Dryopteris hezhangensis P.S.Wang, 2001
Dryopteris himachalensis Fraser-Jenk., 1989
Dryopteris hirtipes (Blume) Kuntze, 1891
Dryopteris × hisatsuana Sa.Kurata, 1964

    (D. handeliana × D. lacera)

Dryopteris × holttumii Li Bing Zhang, 2013

    (D. apiciflora × D. maximowicziana)

Dryopteris hondoensis Koidz., 1932
Dryopteris huberi C.Chr., 1913
Dryopteris huongii C.W.Chen, 2023




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