Le genre Dryopteris 2/4

Source :

344 espèces enregistrées sous le nom

de genre Dryopteris Adans.,

344 espèces légitimes (acceptées),

plus 88 hybrides plus 29 sous-espèces,

18 variétés, 6 formes.

Dryopteris se décline également par des hybridations

entre sous-espèce, variété ou forme.

Ces hybridations plus complexes sont désignés par :

-  6 nothosubsp. ( hybride entre ou avec au moins 1 sous-espèce),

- 5 nothovar. (hybride entre ou  avec au moins 1 variété)

- 1 nothof. (hybride entre ou avec au moins 1 forme)

De A à H

A/
Dryopteris acutissima H.J.Wei & Z.Y.Zuo, 2025

Dryopteris acutodentata Ching, 1938
Dryopteris adnata (Blume) Alderw., 1909
Dryopteris aemula (Aiton) Kuntze, 1891

Dryopteris affinis (Lowe) Fraser-Jenk., 1979, nom. cons.

⤷ affinis subsp. atropes Fraser-Jenk., 2022
⤷ affinis subsp. borreri (Newman) Fraser-Jenk., 1980
⤷ affinis subsp. cambrensis Fraser-Jenk., 1987
⤷ affinis subsp. jessenii (Fraser-Jenk.) Fraser-Jenk., 2007
⤷ affinis subsp. persica Fraser-Jenk., 1980
⤷ affinis subsp. pontica Fraser-Jenk., 1996
⤷ affinis subsp. pseudocomplexa (Fraser-Jenk.) Fraser-Jenk., 2022
⤷ affinis subsp. pseudodisjuncta (Fraser-Jenk.) Fraser-Jenk., 1996
 ⤷ affinis var. austrogallica Fraser-Jenk. & Boudrie, 2024
 ⤷ affinis var. distans (Viv.) Fraser-Jenk., 2024
 ⤷ affinis var. kerryensis Fraser-Jenk., 1996
 ⤷ affinis var. paleaceocrispa (T.Moore) Fraser-Jenk., 2021
 ⤷ affinis var. paleaceolobata (T.Moore) Fraser-Jenk., 2021
 ⤷ affinis var. splendens (Ehrler ex Becherer) Fraser-Jenk., 1980

 ⤷ affinis f. azorica (Fraser-Jenk.) Fraser-Jenk., 2024
 ⤷ affinis f. cluthensis (A.R.Church) Fraser-Jenk., 2024
 ⤷ affinis f. dilatata (Luerss.) Fraser-Jenk., 2024)
 ⤷ affinis f. picoensis Fraser-Jenk., 2024

Dryopteris aitoniana Pic.Serm., 1951
Dryopteris × alejandrei Pérez Carro & Fern.Areces, 2007

   (D. mindshelkensis × D. oreades)

Dryopteris × algonquinensis D.M.Britton, 1975

    (D. fragrans × D. marginalis)

Dryopteris alpestris Tagawa, 1934
Dryopteris × alpirsbachensis Freigang, Zenner, Bujnoch,

  S.Jess. & Magauer, 2017  (D. carthusiana × D. remota)

Dryopteris amblyodonta J.P.Roux, 2004
Dryopteris × ambroseae Fraser-Jenk. & Jermy, 1977

    (D. dilatata × D. expansa)

Dryopteris amurensis (Milde ) Christ, 1909
Dryopteris aneitensis (Hook.) C.Chr., 1905
Dryopteris angustifrons (T.Moore) Kuntze, 1891
Dryopteris angustipalea Darnaedi, M.Kato & K.Iwats., 1989
Dryopteris annamensis (Tagawa) Li Bing Zhang, 2012
Dryopteris antarctica (Baker) C.Chr., 1934
Dryopteris anthracinisquama Miyam., 1984
Dryopteris apiciflora (Wall. ex Mett.) Kuntze, 1891
Dryopteris approximata Sledge, 1973
Dryopteris × apuana Gibby, S.Jess. & Marchetti, 1996

    (D. mindshelkensis × D. villarii)

Dryopteris aquilinoides (Desv.) C.Chr., 1905
Dryopteris arborescens (Baker) Kuntze, 1891
Dryopteris ardechensis Fraser-Jenk., 1981
Dryopteris × arecesiae Pérez Carro & T.E.Díaz, 1990

    (D. aemula × D. corleyi)

Dryopteris arguta (Kaulf.) Watt, 1867
Dryopteris arunachalensis (Fraser-Jenk. & Benniamin)

  R.Kr.Singh & V.K.Rawat, 2024
Dryopteris ascensionis (Hook.) Kuntze, 1891
Dryopteris assamensis (C.Hope) C.Chr. & Ching, 1933
Dryopteris × asturiensis Fraser-Jenk. & Gibby, 1986

    (D. corleyi × D. pseudomas)

Dryopteris athamantica (Kunze) Kuntze, 1891
Dryopteris atrata (Wall. ex Kunze) Ching, 1933
Dryopteris aurantiaca J.P.Roux, 2005
Dryopteris × australis (Wherry) Small, 1938

    (D. celsa × D. ludoviciana)

Dryopteris austroindica Fraser-Jenk., 1989

B/

Dryopteris bamleriana Rosenst., 1912

Dryopteris baniensis Rosenst., 1909
Dryopteris barbigera (Hook.) Kuntze, 1891
Dryopteris basisora Christ, 1909
Dryopteris × benedictii (Farw.) Wherry, 1961

    (D. carthusiana × D. clintoniana)

Dryopteris bernieri Tardieu, 1956
Dryopteris bhutanica (Fraser-Jenk. & Pariyar) Fraser-Jenk., 2023
Dryopteris bissetiana (Baker) C.Chr., 1905
Dryopteris blanfordii (C.Hope) C.Chr., 1905

⤷ blanfordii subsp. nigrosquamosa (Ching) Fraser-Jenk., 1989

Dryopteris bodinieri (Christ) C.Chr., 1905
Dryopteris bojeri Kuntze, 1891
Dryopteris × boottii (Tuck.) Underw., 1893

    (D. cristata × D. intermedia)

Dryopteris borbasii Litard., 1910
Dryopteris × brathaica Fraser-Jenk. & Reichst., 1977

    (D. carthusiana × D. filix-mas)

Dryopteris × burgessii B.Boivin, 1966

    (D. clintoniana × D. marginalis)

Dryopteris burnatii Christ & Wilczek, 1913

C/

Dryopteris cacaina Tagawa, 1937
Dryopteris campyloptera Clarkson, 1930
Dryopteris camusiae Fraser-Jenk., 1997
Dryopteris × cantabrica Alejandre, Pérez Carro & Fern.Areces,

  2016          (D. corleyi × D. guanchica)
Dryopteris caperata J.P.Roux, 2005

Dryopteris caroli-hopei Fraser-Jenk., 1989
Dryopteris carpatica S.Jess. & Bujnoch, 2022
Dryopteris carthusiana (Vill.) H.P.Fuchs, 1959

 ⤷ carthusiana var. enigmatica Harmaja, 2012

Dryopteris caucasica (A.Braun) Fraser-Jenk. & Corley, 1972
Dryopteris caudipinna Nakai, 1931
Dryopteris × cedroensis Gibby & Widén, 1983

    (D. corleyi × D. guanchica)

Dryopteris celsa (W.Palmer) Knowlt., T.S.Palmer & Pollard ex Small,

  1900
Dryopteris chaerophyllifolia (Zippel) C.Chr., 1905
Dryopteris championii (Benth.) C.Chr., 1933
Dryopteris chichisimensis Nakai ex H.Itô, 1935
Dryopteris chinensis (Baker) Koidz., 1930
Dryopteris christenseniae (Ching) Li Bing Zhang, 2012
Dryopteris chrysocoma (Christ) C.Chr., 1905
Dryopteris cicatricata J.P.Roux, 2005
Dryopteris cinnamomea (Cav.) C.Chr., 1911
Dryopteris clarkei (Baker) Kuntze, 1891
Dryopteris clintoniana (D.C.Eaton) Dowell, 1906
Dryopteris cochleata (D.Don) C.Chr., 1905
Dryopteris cognata (C.Presl) Kuntze, 1891
Dryopteris collucata Charit., 2022
Dryopteris commixta Tagawa, 1933
Dryopteris comorensis (Tardieu) Fraser-Jenk., 1994
Dryopteris × complexa Fraser-Jenk., 1987

    (D. affinis × D. filix-mas)

⤷ × complexa nothosubsp. complanata (Fraser-Jenk.) Fraser-Jenk.,
   2024      (D. affinis subsp. pseudodisjuncta × D. filix-mas)

⤷ × complexa nothosubsp. convoluta (Fraser-Jenk.) Fraser-Jenk.,

   2024       (D. affinis subsp. cambrensis × D. filix-mas)

⤷ × complexa nothosubsp. critica Fraser-Jenk., 1987

   (D. affinis subsp. borreri × D. filix-mas)
⤷ × complexa nothosubsp. inconspicua (Fraser-Jenk.) Fraser-Jenk.,

   2022        (D. affinis subsp. pseudocomplexa × D. filix-mas)
⤷ × complexa nothosubsp. transsilvanica Tlałka, S.Jess., A.Rostański

   & Rojek, 2019       (D. affinis subsp. jessenii × D. filix-mas)

⤷ × complexa nothovar. contorta (Fraser-Jenk.) Fraser-Jenk., 2024

   (D. affinis var. paleaceolobata × D. filix-mas)

⤷ × complexa nothovar. eschelmuelleri (Fraser-Jenk.) Fraser-Jenk. &

   Parris, 2020 pub. 2021   (D. affinis var. splendens × D. filix-mas)

⤷ × complexa nothovar. hibernica Fraser-Jenk. & Trewren, 2024

   (D. affinis var. kerryensis × D. filix-mas)

⤷ × complexa nothovar. occidentalis (Fraser-Jenk.) Fraser-Jenk., 2024
   (D. affinis var. paleaceocrispa × D. filix-mas)
⤷ × complexa nothovar. rufosquama Fraser-Jenk., 2024
   (D. affinis var. insubrica × D. filix-mas)

⤷ × complexa nothof. urbachensis (S.Jess. & Bujnoch) Fraser-Jenk.,
   2024    (D. affinis f. dilatata × D. filix-mas)

Dryopteris conjugata Ching, 1941
Dryopteris conversa Alderw., 1915
Dryopteris coreanomontana Nakai, 1921
Dryopteris corleyi Fraser-Jenk., 
Dryopteris × correllii W.H.Wagner & A.V.Gilman, 2001

    (D. carthusiana × D. goldieana)

Dryopteris costalisora Tagawa, 1934
Dryopteris crassirhizoma Nakai, 1920
Dryopteris crinalis (Hook. & Arn.) C.Chr., 1905

 ⤷ crinalis var. podosora (W.H.Wagner) D.D.Palmer, 2002

Dryopteris crispifolia Rasbach, Reichst. & G.Vida, 1977
Dryopteris cristata (L.) A.Gray, 1848
Dryopteris cycadina (Franch. & Sav.) C.Chr., 1905
Dryopteris cyclopeltidiformis C.Chr., 1937
Dryopteris cyclosorus Alderw., 1924

D/

Dryopteris damingshanensis Li Bing Zhang & H.M.Liu, 2014
Dryopteris daozhenensis P.S.Wang & X.Y.Wang, 2001
Dryopteris decipiens (Hook.) Kuntze, 1891
Dryopteris dehuaensis Ching & K.H.Shing, 1982
Dryopteris deparioides (T.Moore) Kuntze, 1891

⤷ deparioides subsp. ambigua (Sledge) Fraser-Jenk., 1989
⤷ deparioides subsp. concinna C.Chr., 1934

Dryopteris devriesei Rosenst., 1917
Dryopteris × deweveri (Jansen) Jansen & Wachter, 1934

    (D. carthusiana × D. dilatata)

Dryopteris diacalpioides (Ching) Li Bing Zhang, 2012
Dryopteris dibangensis C.Chanda & Fraser-Jenk., 2024

Dryopteris dickinsii (Franch. & Sav.) C.Chr., 1905
Dryopteris dicksonioides (Mett.) Copel., 1932
Dryopteris diffracta (Baker) C.Chr., 1905
Dryopteris dilatata (Hoffm.) A.Gray, 1848

⤷ dilatata subsp. lusitanica Fraser-Jenk., Andreis & Vasco Silva, 2021

Dryopteris × doeppii Rothm., 1945

    (D. affinis × A. carthusiana)

Dryopteris × doluchanovii Askerov, 1979

    (D. dilatata × D. pseudomas)

Dryopteris × dowellii (Farw.) Wherry, 1961

    (D. clintoniana × D. intermedia)

Dryopteris dracomontana Schelpe & N.C.Anthony, 1982
Dryopteris drakei C.Chr., 1905
Dryopteris dulongensis (S.K.Wu & X.Cheng) Li Bing Zhang, 2012

E/
Dryopteris edwardsii Fraser-Jenk., 1997
Dryopteris enneaphylla (Baker) C.Chr., 1905
Dryopteris erythrosora (D.C.Eaton) Kuntze, 1891
Dryopteris erythrovaria K.Hori & N.Murak., 2018
Dryopteris × euxinensis Fraser-Jenk. & Corley, 1972

    (D. caucasica × D. filix-mas)

Dryopteris expansa (C.Presl) Fraser-Jenk. & Jermy, 1977

 ⤷ expansa var. alpina (T.Moore) Viane, 1986
 ⤷ expansa var. willeana (Lid) Elven, 1996

F/
Dryopteris fadenii Pic.Serm., 1984
Dryopteris fangii Ching, Fraser-Jenk. & Z.R.Wang, 1985
Dryopteris fengiana (Ching) ined.
Dryopteris ferruginea (Baker) Kuntze, 1891
Dryopteris filipaleata J.P.Roux, 2004
Dryopteris filix-mas (L.) Schott, 1834

⤷ filix-mas subsp. brittonii Fraser-Jenk. & Widén, 2006
⤷ filix-mas subsp. pseudorigida (Christ) Askerov, 1983

Dryopteris flaccisquama A.Rojas, 2001
Dryopteris flemingii Fraser-Jenk., 1989
Dryopteris formosana (Christ) C.Chr., 1905
Dryopteris fragrans (L.) Schott, 1834
Dryopteris fragrantiformis Tzvelev, 1989
Dryopteris × fraser-jenkinsii Gibby & Widén, 1983

    (D. affinis × D. guanchica

Dryopteris fructuosa (Christ) C.Chr., 1905
Dryopteris × fujipedis Sa.Kurata, 1965

    (D. crassirhizoma × D. lacera)

Dryopteris × furadensis Bennert, Rasbach, K.Rasbach & Viane, 1991

    (D. aitoniana × D. intermedia subsp. maderensis)

Dryopteris fuscipes C.Chr., 1917

Dryopteris fuscoatra (Hillebr.) W.J.Rob., 1912

 ⤷ fuscoatra var. lamoureuxii Fraser-Jenk., 1994

Dryopteris futura A.R.Sm., 1975

G/
Dryopteris gamblei (C.Hope) C.Chr., 1905
Dryopteris gaoligongensis Z.Y.Zuo, J.Mei Lu & D.Z.Li,

  2021 publ. 2022 
Dryopteris gemmifera S.Y.Dong, 2010
Dryopteris × ghatakii Fraser-Jenk., 1989

    (D. cochleata × D. sparsa subsp. rectipinnula)

Dryopteris glabra (Brack.) Kuntze, 1891

 ⤷ glabra var. alboviridis (W.H.Wagner) D.D.Palmer, 2002
 ⤷ glabra var. flynnii D.D.Palmer, 2002
 ⤷ glabra var. hobdyana (W.H.Wagner) D.D.Palmer, 2002

 ⤷ glabra var. nuda (Underw. ex A.Heller) Fraser-Jenk., 1994
 ⤷ glabra var. pusilla (Hillebr.) Fraser-Jenk., 1994

 ⤷ glabra var. soripes (Hillebr.) Herat ex Fraser-Jenk., 1994

Dryopteris glandulosopaleata J.P.Roux, 2004
Dryopteris goeringiana (Kunze) Koidz., 1929
Dryopteris goldieana (Hook. ex Goldie) A.Gray, 1848
Dryopteris × gomerica Gibby & Widén, 1983

    (D. aemula × D. guanchica)

Dryopteris gonggaensis H.S.Kung, Li Bing Zhang & X.S.Guo, 1995
Dryopteris gorgonea J.P.Roux, 2004
Dryopteris × gotenbaensis Nakaike, 1969

    (D. hondoensis × D. uniformis)

Dryopteris × graeca Fraser-Jenk. & Gibby, 1980

    (D. mindshelkensis × D. pallida)

Dryopteris grandifrons Li Bing Zhang, 2012
Dryopteris guanchica Gibby & Jermy, 1977
Dryopteris guangxiensis S.G.Lu, 1999
Dryopteris gymnophylla (Baker) C.Chr., 1905
Dryopteris gymnosora (Makino) C.Chr., 1905

H/
Dryopteris habaensis Ching, 1941
Dryopteris hadanoi Sa.Kurata, 1966
Dryopteris × haganecola Sa.Kurata, 1958

    (D. cycadina × D. uniformis)

Dryopteris × hakonecola Sa.Kurata, 1958

    (D. dickinsii × D. uniformis)

Dryopteris handeliana C.Chr., 1937
Dryopteris hangchowensis Ching, 1938
Dryopteris hasseltii (Blume) C.Chr., 1905
Dryopteris hawaiiensis (Hillebr.) W.J.Rob., 1912
Dryopteris hayatae Tagawa, 1932
Dryopteris hendersonii (Bedd.) C.Chr., 1905
Dryopteris herbacea Alderw., 1915
Dryopteris heterolaena C.Chr., 1924
Dryopteris hezhangensis P.S.Wang, 2001
Dryopteris himachalensis Fraser-Jenk., 1989
Dryopteris hirtipes (Blume) Kuntze, 1891
Dryopteris × hisatsuana Sa.Kurata, 1964

    (D. handeliana × D. lacera)

Dryopteris × holttumii Li Bing Zhang, 2013

    (D. apiciflora × D. maximowicziana)

Dryopteris hondoensis Koidz., 1932
Dryopteris huberi C.Chr., 1913
Dryopteris huongii C.W.Chen, 2023




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